In the 1990s, researchers working at the Roslin Institute in Edinburgh, Scotland, performed cloning experiments in collaboration with PPL Therapeutics in Roslin, Scotland, on human coagulation factor IX, a protein.
Keith Henry Stockman Campbell studied embryo growth and cell differentiation during the twentieth and twenty-first centuries in the UK. In 1995, Campbell and his scientific team used cells grown and differentiated in a laboratory to clone sheep for the first time.
The Roslin Institute was established in 1993 in the village of Roslin, Scotland, as an independent research center by the Biotechnology and Biological Sciences Research Council (BBSRC), and as of 2014 is part of the University of Edinburgh in Edinburgh, Scotland.
In 1995 and 1996, researchers at the Roslin Institute in Edinburgh, Scotland, cloned mammals for the first time.